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{{Ant_Infobox|image=File:ThumbLeafcutterAnt.jpg|name=Leafcutter ant|scientific=''Atta cephalotes''|environment=Rainforest|caste=Queen<br/>Minim<br/>Minor<br/>Media<br/>Major|trait=Fungus farming<br/>Taunt<br/>Heal<br/>Resistance buff<br/>Stun|player=Yes}}
{{Ant_Infobox|image=File:Icon_LeafcutterAntMajor.jpg|name=Leafcutter ant|scientific=''Atta cephalotes''|environment=Rainforest|caste=Queen<br/>Minim<br/>Minor<br/>Media<br/>Major|trait=Fungus farming<br/>Taunt<br/>Heal<br/>Resistance buff<br/>Stun|player=Yes}}
Leafcutter ants (''Atta cephalotes'') are playable as complete colonies in story levels, extra levels and custom games.
Leafcutter ants ''(Atta cephalotes)'' are playable as complete colonies in story missions, extra missions and custom games.


<br />
<br />
== '''Abilities'''==
== '''Abilities'''==
Minims are invulnerable and uncontrollable. Each brood tile can spawn and hold a minim when it is built.
Leafcutter ants attack by biting.  


Leafcutter ants use basic attack by biting. You must choose either taunt or stun majors before playing.  
You must choose either taunt or stun majors before playing.  


* Taunt majors can [[Uncategorized Effect|redirect attacks]] onto themselves and [[Healing Effect|heal]]. They can [[Stat Modification|raise their defense]] at low health at level 3.  
* Taunt majors can [[Disrupting Effect|redirect attacks]] onto themselves and [[Healing Effect|heal]]. They can [[Damage Modifying Effect|raise their defense]] at low health at level 3.  


* Stun majors can [[Disrupting Effect|stun]] enemies. They can affect multiple enemies at once and slow them down afterwards at level 3.
* Stun majors can stun enemies. They [[Speed Modifying Effect|slow multiple enemies down]] and [[Damage Modifying Effect|make them take more damage]] at level 3.


You can unlock [[Formicarium Adaptation|adaptations]] for leafcutter ant majors using Royal Jelly in the main story.
Minims are uncontrollable and invulnerable. They can't attack enemies. Each brood tile can spawn a minim without requiring food when it is built.


<br/>
<br/>
=='''Food processing'''==
=='''Food processing'''==
Leafcutter ants automatically bring leaves from harvestable plants on the surface back to leaf storage. Enemies aren't edible and disappear after 3 minutes.
Leafcutter ants can only feed on fungus.  


Minors slow down while carrying leaves. Mediae and majors don't. Gathered leaves are placed in leaf storage. Each leaf storage tile can hold up to 10/ 40/ 100 leaves.
Leafcutter ants have to gather leaves from harvestable plants on the surface. Leaf storage tiles can hold 10/ 40/ 100 leaves.


Minims bring leaves to brood chambers to grow fungus over time. Each brood tile can hold 20/ 30/ 40 food or waste and cultivate 1 food every second.  
Minims bring leaves to brood chambers to grow fungus over time. Brood tiles can hold 20/ 30/ 40 food or waste and cultivate 1 food every second. Queen tiles can hold fungus but not leaves.


Fungus turns into waste that occupies brood tiles when they are used. Minims need to bring waste to waste storage so that fungus has space to grow. Each waste storage tile can hold 10/ 40/ 100 waste and dissolve 1 waste every 5/ 4/ 2.5 seconds.
Fungus turns into waste that occupies brood tiles when they are used. Minims need to bring waste to waste storage so that fungus has space to grow. Waste storage tiles can hold 10/ 40/ 100 waste and dissolve 1 waste every 5/ 4/ 2.5 seconds.


Leafcutter ants are permanently poisoned after going through occupied waste storage in their nest. Affected ants slow down by 35% and lose 50% of their health every 60 seconds. Minims are immune.
Leafcutter ants are poisoned for 60 seconds after moving on occupied waste storage tiles in their nest. Affected ants move 30% more slowly, attack 50% more slowly and lose 0.83% of their health every second. Effect duration is reset each time the affected ants move on another waste storage tile afterward. Minims are immune.


Waste storage tiles remove 2 upgrade points from adjacent tiles and 1 upgrade point from other tiles which are one-tile away. Highway tiles aren't affected. You can only delete a waste storage tile when it is empty.
Waste storage tiles remove 2 upgrade points from adjacent tiles and 1 upgrade point from other tiles which are one-tile away. Highway tiles aren't affected. You can only delete a waste storage tile when it is empty.
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Waste storage usually can't be completely filled up. The extra space is reserved for the waste carried by the minims. Waste can't be dropped when waste storage become full suddenly.
Waste storage usually can't be completely filled up. The extra space is reserved for the waste carried by the minims. Waste can't be dropped when waste storage become full suddenly.


Each storage tile costs 5 food to build. The first leaf storage tile and the first waste storage tile are free.
The first leaf storage tile and the first waste storage tile are free. Later storage tiles cost 5 food to build.  


You receive 66% of the cost as leaves after deleting a tile. Fungus can only exist on brood tiles so leaves are left on the ground instead.
You receive 66.67% of the building cost as leaves from deleting tiles. Fungus can only exist on brood tiles so leaves are left on the ground instead.


Minims panic and stop working when the queen takes damage. You can keep them calm by preventing the respective pheromone groups from attacking.
Minims panic and stop working when the queen takes damage. You can keep them working by preventing the respective pheromone groups from attacking.


<br />
<br />
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*'''[[New Year's Feast]]''' ''(taunt)''
*'''[[New Year's Feast]]''' ''(taunt)''


You can also encounter non-player leafcutter ants in:  
You can also encounter leafcutter ants in:  
*'''[[The Other Foot]]'''
*'''[[The Other Foot]]'''


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*'''[[Adventure]]'''
*'''[[Adventure]]'''
*'''[[Firebreak]]'''
*'''[[No Ant's Land]]'''


<br/>
<br/>
== '''Notable stats'''==
== '''Notable stats'''==
{|class="mw-collapsible wikitable"
{|class="mw-collapsible wikitable"
|colspan="3"|<big>'''Leafcutter ant queen'''</big>
|colspan="3"|[[File:Icon_LeafcutterAntQueen.jpg|50px]] <big>'''Leafcutter ant queen'''</big>
|-
|-
|width="15%"|'''Common stats'''
|width="15%"|'''Common stats'''
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|
|
*Immune to stun/ confusion/ fleeing
*Immune to stun/ confusion/ fleeing
*Immune to pickup/ knockback
*Immune to pickup (strong)/ knockback (strong)
*Immune to execution
*Immune to execution
|}
|}
 
<br/>
<br/>
{|class="mw-collapsible wikitable"
{|class="mw-collapsible wikitable"
|colspan="3"|<big>'''Leafcutter ant minim'''</big>
|colspan="3"|[[File:Icon_LeafcutterAntMinor.jpg|50px]] <big>'''Leafcutter ant minim'''</big>
|-
|-
|width="15%"|'''Common stats'''
|width="15%"|'''Common stats'''
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<br/>
<br/>
{|class="mw-collapsible wikitable"
{|class="mw-collapsible wikitable"
|colspan="3"|[[File:Leafcutterminor.png]] <big>'''Leafcutter ant minor'''</big>
|colspan="3"|[[File:Icon_LeafcutterAntMinor.jpg|50px]] <big>'''Leafcutter ant minor'''</big> [[File:Icon_Worker.png|25px]]
|-
|-
|width="15%t"|'''Common stats'''
|width="15%t"|'''Common stats'''
|colspan="2"|
|colspan="2"|
*''Cost:'' 20/ 2
*''Cost:'' 20 [2]
*''Health:'' 25/ 35/ 45
*''Health:'' 25/ 35/ 45
*''Movement speed:'' 300/ 340/ 380
*''Movement speed:'' 300/ 340/ 380
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*''Healing speed:'' 0.1/ 0.1/ 0.1
*''Healing speed:'' 0.1/ 0.1/ 0.1
*''Vision radius:'' 400/ 400/ 400
*''Vision radius:'' 400/ 400/ 400
*Encumbered
|-
|-
|'''Damage modifications'''
|'''Damage modifications'''
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*None
*None
|-
|-
|'''Basic attack'''
|'''Attack'''
|width="40%"|
|width="40%"|
*''Damage (physical):'' 0.6/ 0.8/ 1
*''Damage (physical):'' 0.6/ 0.8/ 1
*''Attack speed:'' 1/ 1/ 1
*''Attack speed:'' 1.5/ 1.5/ 1.5
|width="45%"|
|width="45%"|
|}
|}
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<br/>
<br/>
{|class="mw-collapsible wikitable"
{|class="mw-collapsible wikitable"
|colspan="3"|[[File:Leafcuttermedia.png]] <big>'''Leafcutter ant media'''</big>
|colspan="3"|[[File:Icon_LeafcutterAntMedia.jpg|50px]] <big>'''Leafcutter ant media'''</big> [[File:Icon_Melee.png|25px]]
|-
|colspan="3"|Stats are different between <font color="red">story mode</font> and <font color="green">custom game mode</font>.
|-
|-
|width="15%"|'''Common stats'''
|width="15%"|'''Common stats'''
|colspan="2"|
|colspan="2"|
*''Cost:'' 80/ 6
*<font color="red">''Cost:'' 80 [6]</font>
*''Health:'' 100/ 115/ 130
*<font color="green">''Cost:'' 65 [6]</font>
*<font color="red">''Health:'' 100/ 115/ 130</font>
*<font color="green">''Health:'' 80/ 95/ 110</font>
*''Movement speed:'' 310/ 320/ 330
*''Movement speed:'' 310/ 320/ 330
*''Score:'' 14/ 16/ 18
*''Score:'' 14/ 16/ 18
*''Harvest amount:'' 15/ 17/ 20 
*<font color="red">''Harvest amount:'' 17/ 19/ 22</font>
*''Healing speed:'' 0.1/ 0.2/ 0.3  
*<font color="green">''Harvest amount:'' 20/ 24/ 30</font>
*''Healing speed:'' 0.1/ 0.2/ 0.3
*''Vision radius:'' 400/ 400/ 400
*''Vision radius:'' 400/ 400/ 400
|-
|-
|'''Damage modifications'''
|'''Damage modifications'''
|colspan="2"|
|colspan="2"|
*''Armor:'' 0.2/ 0.4/ 0.8
*''Armor:'' 0.2/ 0.4/ 0.8
*''Piercing:'' 20%/ 30%/ 40%
*''Piercing:'' 20%/ 30%/ 40%
|-
|-
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*None
*None
|-
|-
|'''Basic attack'''
|'''Attack'''
|width="40%"|
|width="40%"|
*''Damage (physical):'' 4.5/ 6/ 7
*<font color="red">''Damage (physical):'' 4.5/ 6/ 7</font>
*''Attack speed:'' 1/ 1/ 1.3
*<font color="green">''Damage (physical):'' 4/ 5/ 6</font>
*''Attack speed:'' 1.5/ 1.5/ 2
|width="45%"|
|width="45%"|
|}
|}
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<br />
<br />
{|class="mw-collapsible wikitable"
{|class="mw-collapsible wikitable"
|colspan="3"|[[File:Taunt.png]] <big>'''Leafcutter ant major (taunt)'''</big>
|colspan="3"|[[File:Icon_LeafcutterAntMajor.jpg|50px]] <big>'''Leafcutter ant major (taunt)'''</big> [[File:Icon_Tank.png|25px]]
|-
|-
|width="15%"|'''Common stats'''
|width="15%"|'''Common stats'''
|colspan="2"|
|colspan="2"|
*''Cost:'' 150/ 15
*''Cost:'' 150 [15]
*''Health:'' 200/ 240/ 400
*''Health:'' 200/ 240/ 400
*''Movement speed:'' 300/ 300/ 300
*''Movement speed:'' 300/ 300/ 300
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|'''Immunities'''
|'''Immunities'''
|colspan="2"|
|colspan="2"|
*Immune to pickup/ knockback
*Immune to pickup (weak)/ knockback (weak)
|-
|-
|'''Basic attack'''
|'''Attack'''
|width="40%"|
|width="40%"|
*''Damage (physical):'' 12/ 15/ 22
*''Damage (physical):'' 12/ 15/ 22
*''Attack speed:'' 0.8/ 0.8/ 1
*''Attack speed:'' 1.2/ 1.2/ 1.5
|width="45%"|  
|width="45%"|
|-
|-
|rowspan="2"|'''Taunt'''
|rowspan="2"|'''Taunt'''
|rowspan="2"|
|rowspan="2"|
*''Trigger:'' User has above 30% HP.
*''Trigger:'' User has above 30% HP and isn't mounting.
*''Chance:'' 30%/ 30%/ 30%
*''Chance:'' 30%/ 30%/ 30%
*''Damage (physical):'' 1/ 1/ 1
*''Damage (physical):'' 1/ 1/ 1
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|-
|-
|
|
:'''Heal'''
:'''Healing'''
:*''Total health:'' 20/ 25/ 30
:*''Health per interval:'' 1/ 1/ 1
:*''Interval:'' 0.2/ 0.2/ 0.2
:*''Duration:'' 4/ 5/ 6
:*''Duration:'' 4/ 5/ 6
|-
|-
|'''Buff'''
|'''Boost'''
|
|
*''Trigger:'' User has below 40% HP.
*''Trigger:'' User has below 40% HP and isn't mounting.
*''Cooldown:'' --/ --/ 15
*''Cooldown:'' --/ --/ 15
|
|
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<br/>
<br/>
{|class="mw-collapsible wikitable"
{|class="mw-collapsible wikitable"
|colspan="3"|[[File:Stun.png]] <big>'''Leafcutter ant major (stun)'''</big>
|colspan="3"|[[File:Icon_LeafcutterAntMajor.jpg|50px]] <big>'''Leafcutter ant major (stun)'''</big> [[File:Icon_Tank.png|25px]]
|-
|-
|width="15%"|'''Common stats'''
|width="15%"|'''Common stats'''
|colspan="2"|
|colspan="2"|
*''Cost:'' 150/ 15
*''Cost:'' 150 [15]
*''Health:'' 200/ 240/ 400
*''Health:'' 200/ 240/ 400
*''Movement speed:'' 300/ 300/ 300
*''Movement speed:'' 300/ 300/ 300
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|'''Immunities'''
|'''Immunities'''
|colspan="2"|
|colspan="2"|
*Immune to pickup/ knockback
*Immune to pickup (weak)/ knockback (weak)
|-
|-
|'''Basic attack'''
|'''Attack'''
|width="40%"|
|width="40%"|
*''Damage (physical):'' 12/ 15/ 22
*''Damage (physical):'' 12/ 15/ 22
*''Attack speed:'' 0.8/ 0.8/ 1
*''Attack speed:'' 1.2/ 1.2/ 1.5
|width="45%"|  
|width="45%"|
|-
|-
|rowspan="2"|'''Stun'''
|rowspan="3"|'''Stun'''
|rowspan="2"|
|rowspan="3"|
*''Trigger:'' User isn't mounting.
*''Chance:'' 50%/ 50%/ 50%
*''Chance:'' 50%/ 50%/ 50%
*''Damage (physical):'' 1/ 1/ 1
*''Damage (physical):'' 1/ 1/ 1
Line 280: Line 293:
:'''Speed debuff'''
:'''Speed debuff'''
:*''Movement speed:'' --/ --/ - 90%
:*''Movement speed:'' --/ --/ - 90%
:*''Duration:'' --/ --/ 8
|-
|
:'''Resistance debuff'''
:*''Venom resistance:'' --/ --/ - 30%
:*''Duration:'' --/ --/ 8
:*''Duration:'' --/ --/ 8
|}
|}
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<br/>
<br/>
== '''Trivia''' ==
== '''Trivia''' ==
* Dead leafcutter ants can often be seen in the waste storage. In real life, this is a normal behavior in most ant species because waste storage can also serve as a graveyards.
*Leafcutter ant colonies have nineteen-tiled queen chambers in custom games.


* Leafcutter ant larvae don't spin cocoon. Their naked pupae can be seen in the game.
*You can often see dead leafcutter ants in occupied waste storage.


* Level 3 mediae and majors have 100% chance to fend off [[Phorid Fly|phorid flies]], even if they are carrying leaves. This is represented by two minims hanging on the carried leaves, defending the carriers. This behavior also occurs in real life.
*Leafcutter ants are the only ants that have naked pupae in the game.


* Leafcutter ant colonies start with 19-tiled queen chamber in custom games.
*In real life, minims hang onto leaves carried by other ants to help them fend off [[Phorid Fly|phorid flies]]. In the game, only level 3 mediae and majors can get this protection.


<br />
<br/>
[[Category:Ants]]
[[Category:Eusocial Insects]]
[[Category:Rainforest]]
[[Category:Rainforest]]

Latest revision as of 12:39, 1 August 2026

Information
Common name Leafcutter ant
Scientific name Atta cephalotes
Environment Rainforest
Castes Queen
Minim
Minor
Media
Major
Traits Fungus farming
Taunt
Heal
Resistance buff
Stun
Playable colony Yes


Leafcutter ants (Atta cephalotes) are playable as complete colonies in story missions, extra missions and custom games.


Abilities

Leafcutter ants attack by biting.

You must choose either taunt or stun majors before playing.

Minims are uncontrollable and invulnerable. They can't attack enemies. Each brood tile can spawn a minim without requiring food when it is built.


Food processing

Leafcutter ants can only feed on fungus.

Leafcutter ants have to gather leaves from harvestable plants on the surface. Leaf storage tiles can hold 10/ 40/ 100 leaves.

Minims bring leaves to brood chambers to grow fungus over time. Brood tiles can hold 20/ 30/ 40 food or waste and cultivate 1 food every second. Queen tiles can hold fungus but not leaves.

Fungus turns into waste that occupies brood tiles when they are used. Minims need to bring waste to waste storage so that fungus has space to grow. Waste storage tiles can hold 10/ 40/ 100 waste and dissolve 1 waste every 5/ 4/ 2.5 seconds.

Leafcutter ants are poisoned for 60 seconds after moving on occupied waste storage tiles in their nest. Affected ants move 30% more slowly, attack 50% more slowly and lose 0.83% of their health every second. Effect duration is reset each time the affected ants move on another waste storage tile afterward. Minims are immune.

Waste storage tiles remove 2 upgrade points from adjacent tiles and 1 upgrade point from other tiles which are one-tile away. Highway tiles aren't affected. You can only delete a waste storage tile when it is empty.

Waste storage usually can't be completely filled up. The extra space is reserved for the waste carried by the minims. Waste can't be dropped when waste storage become full suddenly.

The first leaf storage tile and the first waste storage tile are free. Later storage tiles cost 5 food to build.

You receive 66.67% of the building cost as leaves from deleting tiles. Fungus can only exist on brood tiles so leaves are left on the ground instead.

Minims panic and stop working when the queen takes damage. You can keep them working by preventing the respective pheromone groups from attacking.


Appearances

You can play a a leafcutter ant colony in:

You can play with leafcutter ant mediae in:

  • Main story

You can play with leafcutter ant majors in:

  • Main story (either)
  • Extremis (either)
  • Tug of War (taunt)

You can also encounter leafcutter ants in:


Notable stats

Leafcutter ant queen
Common stats
  • Health: 1200
  • Score: 700
  • Healing speed: 1
Damage modifications
  • Physical resistance: 40%
  • Venom resistance: 40%
  • Maximum taken damage: 25
Immunities
  • Immune to stun/ confusion/ fleeing
  • Immune to pickup (strong)/ knockback (strong)
  • Immune to execution


Leafcutter ant minim
Common stats
  • Health: 1
  • Movement speed: 220
  • Harvest amount: 4
Damage modifications
  • None
Immunities
  • None


Leafcutter ant minor
Common stats
  • Cost: 20 [2]
  • Health: 25/ 35/ 45
  • Movement speed: 300/ 340/ 380
  • Score: 5/ 8/ 12
  • Harvest amount: 7/ 8/ 9
  • Healing speed: 0.1/ 0.1/ 0.1
  • Vision radius: 400/ 400/ 400
  • Encumbered
Damage modifications
  • None
Immunities
  • None
Attack
  • Damage (physical): 0.6/ 0.8/ 1
  • Attack speed: 1.5/ 1.5/ 1.5


Leafcutter ant media
Stats are different between story mode and custom game mode.
Common stats
  • Cost: 80 [6]
  • Cost: 65 [6]
  • Health: 100/ 115/ 130
  • Health: 80/ 95/ 110
  • Movement speed: 310/ 320/ 330
  • Score: 14/ 16/ 18
  • Harvest amount: 17/ 19/ 22
  • Harvest amount: 20/ 24/ 30
  • Healing speed: 0.1/ 0.2/ 0.3
  • Vision radius: 400/ 400/ 400
Damage modifications
  • Armor: 0.2/ 0.4/ 0.8
  • Piercing: 20%/ 30%/ 40%
Immunities
  • None
Attack
  • Damage (physical): 4.5/ 6/ 7
  • Damage (physical): 4/ 5/ 6
  • Attack speed: 1.5/ 1.5/ 2


Leafcutter ant major (taunt)
Common stats
  • Cost: 150 [15]
  • Health: 200/ 240/ 400
  • Movement speed: 300/ 300/ 300
  • Score: 30/ 42/ 54
  • Harvest amount: 16/ 16/ 16
  • Healing speed: 0.1/ 0.15/ 0.2
  • Threat: 0.3/ 0.3/ 0.3
  • Vision radius: 450/ 460/ 470
Damage modifications
  • None
Immunities
  • Immune to pickup (weak)/ knockback (weak)
Attack
  • Damage (physical): 12/ 15/ 22
  • Attack speed: 1.2/ 1.2/ 1.5
Taunt
  • Trigger: User has above 30% HP and isn't mounting.
  • Chance: 30%/ 30%/ 30%
  • Damage (physical): 1/ 1/ 1
  • Cooldown: 10/ 10/ 10
Taunt
  • Radius: 100/ 140/ 180
  • Duration: 4/ 5/ 6
Healing
  • Health per interval: 1/ 1/ 1
  • Interval: 0.2/ 0.2/ 0.2
  • Duration: 4/ 5/ 6
Boost
  • Trigger: User has below 40% HP and isn't mounting.
  • Cooldown: --/ --/ 15
Resistance buff
  • Physical resistance: --/ --/ + 80%
  • Venom resistance: --/ --/ + 80%
  • Duration: --/ --/ 4


Leafcutter ant major (stun)
Common stats
  • Cost: 150 [15]
  • Health: 200/ 240/ 400
  • Movement speed: 300/ 300/ 300
  • Score: 30/ 42/ 54
  • Harvest amount: 16/ 16/ 16
  • Healing speed: 0.1/ 0.15/ 0.2
  • Threat: 0.3/ 0.3/ 0.3
  • Vision radius: 450/ 460/ 470
Damage modifications
  • None
Immunities
  • Immune to pickup (weak)/ knockback (weak)
Attack
  • Damage (physical): 12/ 15/ 22
  • Attack speed: 1.2/ 1.2/ 1.5
Stun
  • Trigger: User isn't mounting.
  • Chance: 50%/ 50%/ 50%
  • Damage (physical): 1/ 1/ 1
  • Radius: --/ --/ 40
  • Cooldown: 8/ 7/ 6
Stun
  • Duration: 3/ 3.5/ 4
Speed debuff
  • Movement speed: --/ --/ - 90%
  • Duration: --/ --/ 8
Resistance debuff
  • Venom resistance: --/ --/ - 30%
  • Duration: --/ --/ 8


Trivia

  • Leafcutter ant colonies have nineteen-tiled queen chambers in custom games.
  • You can often see dead leafcutter ants in occupied waste storage.
  • Leafcutter ants are the only ants that have naked pupae in the game.
  • In real life, minims hang onto leaves carried by other ants to help them fend off phorid flies. In the game, only level 3 mediae and majors can get this protection.


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